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Dallasaurus
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Dallasaurus ("Dallas lizard") is a basal from the of . Along with , Dallasaurus is one of the two oldest mosasauroid currently known from North America. It is also one of the smallest known , measuring approximately in length.


Specimens
The is based upon two partial skeletons recovered from the Arcadia Park Shale (lower Middle ), approximately 15 meters above its contact with the older Kamp Ranch Limestone in Dallas County in north-central .

The specimen (TMM 43209-1, Texas Memorial Museum, University of Texas at Austin) consists of an incomplete and disarticulated skull, along with considerable portions of the postcranial skeleton, making up about 80 percent of the animal. "Dino-Era Lizard Is Missing Link to Swimming Reptiles, Experts Say", National Geographic News, 21 November 2005. The second referred specimen (DMNH 8121-8125, 8143-8149, and 8161-8180, Dallas Museum of Natural History) lacks any skull material and consists entirely of disarticulated postcranial remains. The containing these fossils were temporarily exposed during excavations for a housing development, and both sites have now been reburied by construction. The two specimens were discovered about 100 meters from one another; the first was found by an amateur collector, Van Turner, for whom the type species ( Dallasaurus turneri) was named. "SMU and Dallas Museum of Natural History Announce Missing Fossil Link Dallasaurus" , Southern Methodist University, 16 November 2005. The genus is named for Dallas County, where both specimens were found.


Anatomy
Polcyn and Bell diagnose Dallasaurus as follows: "Small, plesiopedal mosasauroid possessing the following autapomorphies: posterior teeth strongly recurved posteriorly, slightly inflated at the crown and bearing only posterior carinae that is slightly offset laterally; atlas neural arch mediolaterally compressed but not flattened at its base, condylar surfaces irregularly figure-eight shaped; cervical vertebra synapophyses protrude below the level of the edge of the centrum; short, wide fossa excavated immediately below the ventral rim of the cotyle of at least one middle cervical vertebra; hypapophysis anteroventral edge terminating in short projections of irregular length; postglenoid process capped by bony bearing a calcified apex."Bell & Polcyn, Dallasaurus turneri, pp. 179-180.

Bell and Polcyn use the term "plesiopedal" to indicate a "conservative ecologically adaptive grade" characterized by "small size, slightly modified swimming tail and a relatively limb condition" compared to more derived mosasauroids. Polcyn and Bell note that plesiopedal mosasauroids tend to be relatively small lizards possessing limbs in which the "propodial elements equals remain elongated, generally constituting one-half or more of the full length of the osseus limb",Bell & Polcyn, Dallasaurus turneri, p. 178. as compared to more derived "hydropedal" mosasaurs in which the propodial elements are stout and have been substantially shortened, constituting less than one-half of the full length of the ossueus limb. While hydropedal mosasaurs were probably entirely aquatic, plesiopedal mosasauroids were still capable of terrestrial locomotion and so likely lived an amphibious lifestyle.


Classification
Bell and Polcyn have conducted a cladistic analysis of Dallasaurus, concluding that this should be placed within the subfamily , "well within the ", and that, despite its small size and the "primitive" condition of its limbs, it should not be placed within the family .Bell & Polcyn, Dallasaurus turneri, pp. 188-190. Dallasaurus is the to more derived mosasaurine mosasaurs such as , , Mosasaurus, and .

In the popular press, Dallasaurus has been hailed as a "missing link" uniting fully aquatic mosasaurs with their terrestrial ancestors. "Missing fossil link 'Dallasaurus' found", Science Daily, 16 November 2005.


Sources
  • G. L. Bell, Jr.; Polcyn, M. J. (2005). " Dallasaurus turneri, a new primitive mosasauroid from the Middle Turonian of Texas and comments on the phylogeny of the Mosasauridae (Squamata)." Netherlands Journal of Geoscience ( Geologie en Mijnbouw) 84 (3) pp. 177–194.

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